iii

INTRODUCTION TO

GENETIC

ANALYSIS

ELEVENTH EDITION

Anthony J. F. Griffiths

University of British Columbia

Susan R. Wessler

University of California, Riverside

Sean B. Carroll

Howard Hughes Medical Institute

University of Wisconsin–Madison

John Doebley

University of Wisconsin–Madison

iv

Publisher

      Kate Ahr Parker

Senior Acquisitions Editor

      Lauren Schultz

Executive Marketing Manager

      John Britch

Marketing Assistant

      Bailey James

Developmental Editor

      Erica Champion

Editorial Assistant

      Alexandra Garrett

Supplements Editor

      Erica Champion

Executive Media Editor

      Amanda Dunning

Media Editors

      Donna Broadman, Erica Champion, Tue Tran

Art Director

      Diana Blume

Cover and Interior Designer

      Vicki Tomaselli

Senior Project Editor

      Jane O’Neill

Permissions Manager

      Jennifer MacMillan

Photo Editor

      Richard Fox

Illustration Coordinator

      Janice Donnola

Illustrations

      Dragonfly Media Group

Production Supervisor

      Susan Wein

Composition and Layout

      Sheridan Sellers

Printing and Binding

      RR Donnelley

Cover Photo

      Susan Schmitz/Shutterstock

Library of Congress Preassigned Control Number: 2014957104

Hardcover: ISBN-13: 978-1-4641-0948-5

     ISBN-10: 1-4641-0948-6

Loose-Leaf:   ISBN-13: 978-1-4641-8804-6

    ISBN-10: 1-4641-8804-1

© 2015, 2012, 2008, 2005 by W. H. Freeman and Company

All rights reserved

Printed in the United States of America

Fourth printing

W. H. Freeman and Company

A Macmillan Education Imprint

41 Madison Avenue

New York, NY 10010

Houndmills, Basingstoke RG21 6XS, England

www.macmillanhighered.com

v