| YEAR 1 | YEAR 2 | |||||||
|---|---|---|---|---|---|---|---|---|
| MALE ROLE | B1 | r1 | INDIRECT FITNESS | B2 | r2 | Psm | DIRECT FITNESS | INCLUSIVE FITNESS |
| PRIMARY HELPER | 1.0 × 0.32 = 0.32 | 2.5 × 0.5 × 0.32 = 0.41 | 0.32 + 0.41 = 0.73 | |||||
| SECONDARY HELPER | 1.3 × 0.00 = 0.00 | 2.5 × 0.5 × 0.67 = 0.84 | 0.00 + 0.84 = 0.84 | |||||
| DELAYER | 0.0 × 0.00 = 0.00 | 2.5 × 0.5 × 0.23 = 0.29 | 0.00 + 0.29 = 0.29 | |||||