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for the Eleventh Edition

Advisory Board

Lori Ann Biederman, Iowa State University

Candice Damiani, Unversity of Pittsburgh

Claudette Davis, George Mason University, Fairfax

Kamal Dulai, University of California, Merced

Ellen S. Goldey, Florida Atlantic University

Christopher Gregg, Louisiana State University

Carly Jordan, The George Washington University

Brenda Leicht, University of Iowa

Kristen Lennon, Hagerstown Community College

Jay Mager, Ohio Northern University

David J. Marcey, California Lutheran University

Robert Osuna, University at Albany, State University of New York

Pamela A. Pape-Lindstrom, Everett Community College

Roger Persell, Hunter College, City University of New York

Anna Powolny, Spelman College

Mitchell Singer, University of California, Davis

Kathryn Spilios, Boston University

Sarah Texel, Towson University

Catherine Ueckert, Northern Arizona University

Between Edition Reviewers

Elizabeth Binney, Western Washington University

Catherine Black, Idaho State University

Andrew Blaustein, Oregon State University

Katie Boes, College of Wooster

Michelle Boone, Miami University of Ohio

Christopher Brown, Georgia Gwinnett College

Theodore Burk, Creighton University

Cheryl Burrell, Wake Forest University

Jean Cardinale, Alfred University

Shelton Charles, Forsythe Technical Community College

Debora Christensen, Drake University

John Cigliano, Cedar Crest College

Beverly Clendening, Hofstra University

Catherine Craker, Ohio Northern University

Noelle Cutter, Molloy College

Claudette Davis, George Mason University

Amy Downing, Ohio Wesleyan University

William Eldred, Boston University

David Eldridge, Baylor University

Julie Emerson, Amherst College

Stuart Feinstein, University of California, Santa Barbara

Kenneth Filchak, University of Notre Dame

Teresa Fischer, Indian River State College

Pam Freeman, The College of St. Scholastica

Theresa Grove, Valdosta State University

Kathy Hafer, Washington University in St. Louis

Kevin Hamed, Virginia Highlands Community College

Laura L. Hernandez, University of Wisconsin, Madison

Mark Holland, Salisbury University

Lara Hutson, State University of New York at Buffalo

Danielle Ignace, Smith College

David Kittlesen, University of Virginia

Drew Kohlhorst, Georgia State University

Stephen Kolomyjec, Ohio Northern University

Kristen Lennon, Hagerstown Community College

Eric Liebold, Salisbury University

Robert Lowery, Indian River State College

Jay Mager, Ohio Northern University

Stephanie Marin, California State Polytechnic University, Pomona

Elizabeth McCain, Muhlenberg College

Andy McCall, Denison University

Clara Moore, Franklin and Marshall College

Richard Moore, Miami University of Ohio

Ross Nehm, Stony Brook University, State University of New York

Leah Okumura, Wellesley College

Nancy Pencoe, University of West Georgia

Uwe Pott, University of Wisconsin, Green Bay

David Puthoff, Frostburg State University

Diviya Ray, Massachusetts Institute of Technology

Brian Ring, Valdosta State University

Mary K. Ritke, University of Indianapolis

George Robinson, University at Albany, State University of New York

Jason Rothman, California State Polytechnic University, Pomona

Patricia Rugaber, College of Coastal Georgia

Mark Sarvary, Cornell University

Robert Savage, Williams College

Mark Schneegurt, Wichita State University

Fayla Schwartz, Everett Community College

Wayne Silver, Wake Forest University

Neelima Sinha, University of California, Davis

Joel Snodgrass, Towson University

Sita Somara, Forsythe Technical Community College

Kathryn Spilios, Boston University

Cynthia Surmacz, Bloomsburg University

Steven Swoap, Williams College

Edward Tall, Seton Hall University

Mark Taylor, Baylor University

Douglas Thrower, University of California, Santa Barbara

Dean Tolan, Boston University

Catherine Ueckert, Northern Arizona University

Laura Vallier, Hofstra University

John Walker, Appalachian State University

Heping Zhou, Seton Hall University

Accuracy Reviewers

Cristian Aguilar, Azusa Pacific University

C. A. Allen, Blinn University

Tracey Anderson, University of Minnesota, Morris

Caleb Bailey, Brigham Young University, Idaho

Michael Balsai, Temple University

Michael Baltzley, Western Oregon University

Adebiyi Banjoko, Chandler Gilbert Community College

Chad Barber, California Lutheran University

Todd Barkman, Western Michigan University

Alexandra Bely, University of Maryland

Michael Berger, Washington State University, Vancouver

Karen K. Bernd, Davidson College

Stephanie Bingham, Barry University

Elizabeth Binney, Western Washington University

Kristopher Blee, California State University, Chico

Ed Bobich, California State Polytechnic University, Pomona

Robert Bohanan, University of Wisconsin

Nancy Boury, Iowa State University

Melissa Bowlin, University of Michigan, Dearborn

Sarah Boyer, Macalester College

xxviii

Forrest Michael Brem, University of Memphis

Matthew W. Brewer, Georgia State University

Mirjana Brockett, Georgia Institute of Technology

Heather Bruns, Ball State University

Jill Buettner, Richland College

Steve Burnett, Clayton State University

Cheryl Burrell, Forsyth Technical Community College

Steve Bush, Coastal Carolina University

Madhavi Chakravadhanula, Arizona State University

Eric Chambers, Valdosta State University

Shelton Charles, Forsyth Technical Community College

Charles Chen, Azusa Pacific University

Vasanta Chivukula, Atlanta Metro College

Lynn Christenson, Vassar College

Dennis O. Clegg, University of California, Santa Barbara

Craig Clifford, Northeastern State University

Liz Co, Boston University

Reid Compton, University of Maryland

Stephanie Conant, University of Detroit, Mercy

Vikki Connaughton, American University

Cindy Corbitt, University of Louisville

Brett Couch, University of British Columbia

Alistair Cullum, Creighton University

Noelle Cutter, Molloy College

Mary Ellen Czesak, Vassar College

David Dalton, Reed College

Deborah Dardis, Southeastern Louisiana University

Doug Darnowski, Indiana University, Southeast

Farahad Dastoor, University Maine

Andrew David, Clarkson University

Frank David, University of California, Santa Barbara

William Davis, Washington State University

Lewis Deaton, University Louisiana, Lafayette

Kelly Decker, California State University, East Bay

John Demastes, University of Northern Iowa

Fred Deneke, Cosumnes River College

Jill DeVito, University of Texas, Arlington

Sam Donovan, University of Pittsburgh

Amy Downing, Ohio Wesleyan University

Matthew Draud, Armstrong State University

Robert Drewell, Clark University

Kelli Duncan, Vassar College

Erin Easlon, University of California, Davis

John Elder, Valdosta State University

Jean Everett, College of Charleston

Gordon Fain, University of California, Los Angeles

David Fankhauser, University of Cincinnati, Clermont

Melinda Faulkner, Bradley University

Kenneth Filchak, University of Notre Dame

Samuel Flaxman, University of Colorado, Boulder

Amy Frary, Mount Holyoke College

Aaren Freeman, Adelphi University

Mark Garcia, Collin College

Marina Gerson, California State University, Stanislaus

Raffaella Ghittoni, University of Southern California

Brian Gibbens, University of Minnesota

J. Phil Gibson, University of Oklahoma

William Gilliland, DePaul University

Jaime Grace, Bradley University

Matthew Graham, Eastern Connecticut State University

Robert Grammer, Belmont University

Mike Grosbeck, Brigham Young University, Idaho

Theresa Grove, Valdosta State University

Lisa Grubisha, University Wisconsin, Green Bay

Ehren Haderlie, Brigham Young University, Idaho

Rebecca Hale, University of North Carolina at Asheville

Ben Hanelt, University of New Mexico

Amy Hark, Muhlenberg College

Phillip Harris, University of Alabama, Tuscaloosa

Valerie Haywood, Case Western Reserve University

Christiane Healey, University of Massachusetts, Amherst

Tracy Heath, Iowa State University

Heather Heck, University of Maine

Robert Hegna, Palm Beach Atlantic University

Amy Helms, Collin College

Kelly Howe, University of New Mexico

David Hudson, Atlanta Metropolitan State College

Myra Hughey, Vassar College

Ryan Huish, The University of Virginia’s College at Wise

Cynthia Hutton, Northland Pioneer College

Chris Ivey, California State University, Chico

Erika Iyengar, Muhlenberg University

Rachel Johnson, University of Minnesota, Morris

Keith Johnson, Bradley University

Scott Johnson, Towson University

Russell Johnson, Colby College

Jennifer Jost, Bradley University

Jacob Kagey, University of Detroit, Mercy

Stephanie Kamel, University North Carolina, Wilmington

Lori Kayes, Oregon State University

David Keller, California State University, Chico

Diane Kelly, University of Massachusetts, Amherst

Todd Kelson, Brigham Young University Idaho

Kyeorda Kemp, Northeastern State University

Tess Killpack, Wellesley College

Scott Kimball, Baker University

Lara Kingeter, Tarrant County College | Northeast Campus

Bridgette Kirkpatrick, Collin College

David Kittlesen, University of Virginia

David Knochel, University of Colorado, Denver

William Kristan, California State University, San Marco

Allan Larson, Washington University in St. Louis

Kristin Latham, Western Oregon University

Kari Lavalli, Boston University

Mark Lazzaro, College of Charleston

Kay Lee-Fruman, California State University, Long Beach

Jeff Leips, University Maryland, Baltimore County

Kristen Lennon, Hagerstown Community College

Eric Liebl, Denison University

Brianna Lindh, Willamette University

Rob Loeb, Pennsylvania State University

John Logsdon, The University of Iowa

Janet Loxterman, Idaho State University

Charles Mallery, University of Miami, Coral Gables

Robert Mans, Armstong State University

Barry Margulies, Towson University

Linda Mayerhofer, University at Albany, State University of New York

Bryant McAllister, The University of Iowa

Lori McGrew, Belmont University

Kyle McQuade, Colorado Mesa University

Mark Meade, Jacksonville State University

Brad Mehrtens, University of Illinois at Urbana-Champaign

Jennifer Metzler, Ball State University

Rick Miller, Formerly at Southeastern Louisiana University

Rich Moore, Miami University, Ohio

Patricia Moore, University of Georgia

Sherri Morris, Bradley University

Barbara Musolf, Clayton State University

Ross Nehm, Stony Brook University, State University of New York

Kim Nelson, Pennsylvania State University

Katie Nemeth, College of St. Scholastica

John Niedzwiecki, Belmont University

Ben Normark, University of Massachusetts, Amherst

Eric Norstrom, DePaul University

Richard Noyes, University of Central Arkansas

Matthew Nusnbaum, Georgia State University

Leah Okumura, Wellesley College

Jonelle Orridge, Broward College

Jennifer Osterhage, University of Kentucky

Kristen Patterson, University of Texas at Austin

xxix

James Patton, Vanderbilt University

Paulette Peckol, Smith College

Patricia Peroni, Davidson College

Teresa Petrino, Barry University

Uwe Pott, University of Wisconsin-Green Bay

Dan Potter, University of California, Davis

Daniel Potts, Buffalo State University

Nick Pullen, University Northern Colorado

Nick Ragsdale, Belmont University

Claire Varian Ramos, Colorado State University, Pueblo

Flona Redway, Barry University

Melissa Reedy, University of Illinois Urbana-Champaign

Mark Reedy, Creighton University

Susan Reigler, Indiana University Southeast

Laurel Roberts, University of Pittsburgh

George Robinson, University at Albany, State University of New York

Kevin Roe, Iowa State University

Corey Roelke, University Texas at Arlington

Kristi Curry Rogers, Macalester College

Thomas Rosburg, Drake University

Mike Rosenzweig, Virginia Tech

Jason Rothman, California State Polytechnic University, Pomona

Ben Rowley, University of Central Arkansas

Jodi Rymer, College of the Holy Cross

Robert Savage, Williams College

Brenda Schoffstall, Barry University

Paul Schulter, University of Nevada, Las Vegas

J. Reid Schwebach, George Mason University

Erik Scully, Towson University

Jeanne Serb, Iowa State University

Alan Shabel, University of California. Berkeley

Mike Shaughnessy, Northeastern State University

Mark Sherrard, University of Northern Iowa

Ann Showalter, Clayton State University

Cahleen Shrier, Azusa Pacific University

Wayne Silver, Wake Forest University

Anu Singh-Cundy, Western Washington University

Diviya Sinha, Massachusetts Institute of Technology

Bill Smith, Wake Forest University

Nancy Solomon, Miami University, Ohio

Jay Stachowicz, University of California, Davis

Maria Stanko, New Jersey Institute of Technology

Nathan Staples, Canada College

Karl Sternberg, Western New England University

Tara Turley Stoulig, Southeastern Louisiana University

M. Elizabeth Stroupe, Florida State University

Ken Sweat, Arizona State University

Beckie Symula, University of Mississippi

Nicoladie Tam, University of North Texas

Amina Tassa, Collin College

Casey terHorst, California State University, Northridge

Conrad Toepfer, Brescia University

Dean Tolan, Boston University

W. Robert Trentham, Carson Newman University

Claudia Uhde-Stone, California State University, East Bay

Hemayet Ullah, Howard University

Ximena Valderrama, Ramapo College of New Jersey

Kenneth van Golen, University of Delaware

Doug Van Hoewyk, Coastal Carolina University

John VandenBrooks, Midwestern University

Meenakshi Vijayaraghavan, Tulane University

Janice Voltzow, University of Scranton

Catherine Vrentas, Frostburg State University

Heather Waye, University Minnesota, Morris

Allison Welch, College of Charleston

Gerard White, College of Coastal Georgia

Mary White, Southern Louisiana University

Lisa Whitenack, Allegheny College

Judith Williams, University of San Diego

Daniel Williams, Coastal Carolina University

Gay Williamson, Mississippi State University

Sarah Wojiski, Massachusetts College of Pharmacy and Health Sciences

Weimin Zhong, Yale University

Pedagogy Reviewers

Lori Biederman, Iowa State University

Jennifer Butler, Willamette University

Liz Co, Boston University

Julie Cronk, Columbus State Community College

Deborah Dardis, Southeastern Louisiana University

Farahad Dastoor, University of Maine

Claudette Davis, George Mason University

Tod Duncan, University of Colorado, Denver

Miles Dean Engell, North Carolina State University

Ginger Fisher, University of Northern Colorado

Marina Gerson, California State University, Stanislaus

Valerie Haywood, Case Western Reserve University

Ryan Huish, The University of Virginia’s College at Wise

Jamie Jensen, Brigham Young University

Rachel Johnson, University of Minnesota, Morris

Dave Kittlesen, University of Virginia

Kari Lavalli, Boston University

Jeanne Lawless, Binghampton University, State University of New York

Brenda Leicht, University of Iowa

Kristen Lennon, Hagerstown Community College

Carolyn Martineau, DePaul University

Dina Newman, Rochester Institute of Technology

Robert Osuna, University at Albany, State University of New York

Mike Rosenzweig, Virginia Tech

Mitch Singer, University of California, Davis

Alicia Slater, Stetson University

Kathryn Spilios, Boston University

Ximena Valderrama Ramapo, The College of New Jersey

Catherine Wilcoxson Ueckert, Northern Arizona University

Mary Williams, American Society of Plant Biology, Features Editor, The Plant Cell

Robert Wise, University Wisconsin, Osh Kosh

Media and Supplements Contributors

Edward Awad, Vanier College

Susan Lopez Bailey, science writer

Cissy Ballen, Cornell University

Joseph Bruseo, Holyoke Community College

Kenneth Filchak, University of Notre Dame

Donna Francis, University of Massachusetts, Amherst

Richard Gill, Brigham Young University

Richard Gonzalez-Diaz, Seminole State College

Adam Hall, Smith College

Carol Hand, science writer

Phillip Harris, University of Alabama

Margaret Hill, science writer

Norman Johnson, University of Massachusetts Amherst

Carly Jordan, George Washington University

Jessica Kalagher, science writer

Brenda Leicht, University of Iowa

Laurie Beth Leonelli, Sarah Lawrence College

John Lepri, University of North Carolina at Greensboro

Betty McGuire, Cornell University

Polly Molyneux, Smith College

Kristine Nowak, Kennesaw State University

Scott Nowak, Kennesaw State University

Diviya Ray, Massachusetts Institute of Technology

Meredith Safford, Johns Hopkins University

Jessica Santangelo, Hofstra University

John Townsend-Mehler, Montana State University

Mary Tyler, University of Maine

Danielle Webster, University of Central Florida

Robert Wise, University of Wisconsin, Oshkosh

Timothy Wakefield, John Brown University