7.3 Soil structure and fertility result from dynamic processes

Soils are more than just dirt. Plants depend on them for physical support, water, and nutrients. They are also home to diverse underground organisms. Creatures living in soils range from microscopic bacteria to the largest organism known—a honey fungus, Armillaria ostoyae, in eastern Oregon, which is 3 times the size of New York’s Central Park (Figure 7.5). Soil development and soil structure vary around the world largely because of differences in climate and the types and abundance of organisms present (Figure 7.6).

TEEMING WITH LIFE, SOILS ARE HOME TO A VAST ARRAY OF BIODIVERSITY
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FIGURE 7.5 Many thousands of bacterial species can be identified from a single gram of soil. Shown here are just a few of the vast number of life forms that inhabit soils.
(Hugh Spencer/Science Source; Colorization by Mary Martin (Nigel Cattin/Science Source) (D. Kucharski K. Kucharska/Shutterstock) (stshank/Shutterstock)
COMBINED EFFECTS OF CLIMATE AND ORGANISMS PRODUCE MAJOR DIFFERENCES AMONG THE TYPICAL SOILS OF EARTH’S BIOMES
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FIGURE 7.6 Differences in climate and dominant organisms in terrestrial biomes are reflected in variations in amounts of organic matter, depth, color, and fertility of their soils.

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Soil Structure

O horizon The surface layer of many soils, which is rich in organic matter and a site of active decomposition.

If you were to dig into a mature soil, you would find a series of layers, called soil horizons. In a temperate deciduous forest, these are known as the O, A, E, B, C, and R horizons (Figure 7.7).

BASIC SOIL STRUCTURE CONSISTS OF A VERTICAL SEQUENCE OF SOIL HORIZONS
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FIGURE 7.7 Mature soils under temperate deciduous forests commonly have six horizons: the O, A, E, B, C, and R horizons.

The surface layer, or the O horizon, is a site of active decomposition of organic matter, such as leaves, twigs, and bark. Burrowing organisms and physical processes, such as freezing and thawing, mix this decomposing organic matter with the inorganic clays and sands found in the lower soil horizons, producing the crumblike structure characteristic of many fertile soils.

A horizon (topsoil) Soil layer immediately below the O horizon that includes significant amounts of organic matter, generally expressed by dark color.

The A horizon, which sits beneath the the O horizon, is typically referred to as the topsoil. It is rich in essential plant nutrients, such as nitrogen, phosphorus, and potassium, on which much plant production is dependent. Although it contains significant amounts of organic matter, which is generally dark in color, the A horizon is predominantly an inorganic layer, consisting of a mixture of various proportions of sand, silt, and clay.

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Why are soils with a deep A horizon generally considered good for farming?

soil texture The relative fineness or coarseness of a soil, which is determined by its proportions of sand, silt, and clay.

loam A soil consisting of approximately equal proportions of sand, silt, and clay.

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The proportions of sand (coarse and gritty), silt (intermediate grain size), and clay (small grain size) in the topsoil determine soil texture. Soils dominated by one of the three types of mineral soil particles are called sandy, silty, or clay soils. Meanwhile, a soil consisting of approximately equal proportions of sand, silt, and clay is a loam. A loam soil, which has properties intermediate between sandy and clay soils, is considered one of the most desirable for agriculture.

E horizon Soil layer between the A and B horizons, from which clays and dissolved materials are transported down the soil profile to the underlying B horizon.

B horizon A depositional soil layer in which materials transported from the A and E horizons accumulate.

C horizon The deepest soil layer, consisting mainly of lightly weathered parent material.

parent material The bedrock or unconsolidated deposits, such as windblown sand or silt, from which soil develops.

R horizon The base of a soil profile composed of consolidated bedrock, immediately below the C horizon.

In well-developed soils, there can be an E horizon below the A horizon. The light-colored E horizon results from clays and dissolved organic matter flowing downward to deeper soil layers, leaving pale-colored sand and silt. The subsoil, or B horizon, is a depositional layer, rich in materials that leached out of the E horizon. The C horizon, the deepest soil layer, typically contains unconsolidated weathered rocks from the parent material from which soil develops. Parent material can be made up of rock, windblown or water-transported sand, or organic matter, such as peat. At the base of a soil developed on rock is the R horizon, which is partially weathered bedrock.

Soil Development

Soil forms as the environment interacts with parent material. The factors important to soil formation include climate, organisms, the nature of the parent material, the topography (or form) of the land surface, and time (Figure 7.8).

SOIL DEVELOPMENT IS A SLOW PROCESS
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FIGURE 7.8 Soils form as a consequence of climate and organisms acting on parent material over long periods of time. The sequence here sketches soil development in a climate that would support temperate deciduous forest at maturity.

Climate directly influences soil development through temperature and precipitation. Temperature fluctuations, freezing and thawing in cold climates, and heating and cooling in hot climates, promote weathering of rocks. Weathering begins with fracturing and fragmentation of large rock, which eventually reduces even great rock formations to small soil particles. The rate of soil development reaches maximum levels in the warm, humid tropics. Climate also indirectly affects soil development through its influences on the activity of soil organisms and plant roots.

erosion A process that removes geologic materials, ranging from clay-sized particles to boulders, from one part of a landscape to be deposited elsewhere; increased rates of soil erosion due to human activity can reduce soil fertility.

Wind and rain add nutrients as they deposit dust on a landscape (Figure 7.9). Nitrogen-fixing bacteria and plants, such as legumes, produce most of the biologically available nitrogen in soils. Runoff from rainfall can cause erosion of soils from the landscape, whereas rainfall that percolates into soils can remove nutrients and carry them down the soil profile and into groundwater.

SOIL STRUCTURE AND FERTILITY RESULT FROM A DYNAMIC INTERPLAY OF PROCESSES
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FIGURE 7.9 Avenues of soil nutrient loss oppose several sources of nutrient addition. Soil nutrients taken up by plant roots and carbon dioxide from the atmosphere are incorporated into plant tissues during photosynthesis. Then as plant litter and other organic matter decompose (e.g., fallen leaves, dead roots, or shed bark), carbon dioxide is released to the atmosphere and soil nutrients are returned to the soil.

Though all soils are subject to erosion by wind or water, those on steep slopes are more vulnerable. As soils erode from higher ground, the soil remaining on the slopes is thin and prone to drying. Soils washed into valleys and other low points in a landscape, such as swales, cause a thickening and moistening in these depositional areas.

Think About It

  1. Why might some plants grow well on soils in the early stages of development but not in later stages (see Figure 7.8)?

  2. What properties might lead farmers to consider loam to be an ideal soil?

  3. How would differences in the main factors influencing soil development (climate, organisms, parent material, topography, and time) influence the store of nutrients and organic matter in soils as depicted in Figure 7.9?

  4. If global climate changes such that the optimal zone for grain production moves far to the north, how might soils limit agriculture in this new “climatically optimal” zone?

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7.1–7.3 Science: Summary

Climate, which includes temperature and precipitation, is one of the most important factors influencing the amount of biomass an ecosystem produces. Careful experimental research has revealed that biodiversity also has a significant positive influence on productivity, independent of climate. These documented relationships may help in the design of sustainable agricultural systems.

Different climatic conditions support a wide range of biomass production and different soil types. This variation in climate and soils is linked to Earth’s terrestrial biomes, within which human populations have developed systems of farming, ranching, and forestry to harvest primary production for human use.

In a mature temperate deciduous forest, soils have a distinct sequence of layers called the O, A, E, B, C, and R horizons. The main factors important to soil formation include climate, organisms, the nature of the parent material, the topography (or form) of the land surface, and time. The supply of essential plant nutrients (e.g., nitrogen and phosphorus) and organic matter in soils is not static, but rather the result of a dynamic interaction between several processes, such as erosion, deposition, and decomposition.