Five hundred million years ago, the land surface supported little more than a green crust made up of a mixture of photosynthetic and non-photosynthetic microbes. Then, through the process of natural selection, a group of freshwater algae began the transition to survival on land. Fast-forward 465 million years—to the present time—and plants are everywhere. Much of what makes up a plant, including roots, leaves, tree trunks, and seeds, evolved quickly. Within the first quarter of the time since these algal ancestors began to colonize the land, all of these features were present. But one thing was missing: flowers. Flowering plants, also known as angiosperms, do not appear on the scene until the last quarter of this history, and yet today they make up a remarkable 90% of all plant species. Many of the plant groups that diverged before angiosperms appeared are still present, but where they live and grow has been shaped by the evolution of flowering plants.